1940
Born in Belfast, Ireland.
1961
Earns a Bachelor of Science in physiology and chemistry from the University of London.
1964
Earns a Doctor of Philosophy in physiology from the University of Sheffield.
1965
Immigrates to the United States.
1965
Is a research fellow at Harvard University through 1966.
1967
Teaches as an assistant professor of physiology at the University of California Los Angeles until 1970.
1970
Teaches as an associate professor of physiology at the University of California Los Angeles until 1974.
1974
Teaches as a professor of physiology at the University of California Los Angeles and presently maintains this role.
1978
Earns a Doctor of Science in physiology from the University of London.
1982
Serves as a consultant for the National Institutes of Health in Bethesda, MD.
1985
Is a neuroscience investigator for Senator Jacob K. Javits.
2000
Is named a Horace W. Davenport Distinguished Lecturer by the American Physiology Society.
2004
Receives the Janssen Gastroenterology Award from the Janssen/American Physiology Society.
2005
Becomes a fellow of the Ropul Society in the UK.
2006
Becomes a fellow of the of the German Academy of Sciences (Leopoldina).
2012
Receives the Homer W. Smith Award from the American Society of Nephrology.
2013
Is a fellow of the National Academy of Science.
2016
Serves on the external advisory board within the Center for Structure of Membrane Proteins, UCS.